Todiramphus cinnamominus · mitochondrial ND2 · 156 sequences

Nothing lives within two percent of the sihek.

Every kingfisher sequence ever deposited, plotted by how far its ND2 gene has drifted from the Guam kingfisher. The bird sits at the centre. The ring around it is empty — not sparse, empty. This chart inks the emptiness and leaves the data as dust.

Sihek 3
3 5 10 15 20 25 ND2 % DIVERGENCE Actenoides bougainvillei excelsus -- Wikipedia: Moustached kingfisher Actenoides concretus -- Wikipedia: Rufous-collared kingfisher Actenoides hombroni -- Wikipedia: Blue-capped kingfisher Actenoides lindsayi -- Wikipedia: Spotted wood kingfisher Caridonax fulgidus -- Wikipedia: White-rumped kingfisher Actenoides monachus -- Wikipedia: Green-backed kingfisher Actenoides princeps -- Wikipedia: Scaly-breasted kingfisher Todiramphus ruficollaris -- Wikipedia: Mewing kingfisher Todiramphus ruficollaris -- Wikipedia: Mewing kingfisher Todiramphus gertrudae -- Wikipedia: Niau kingfisher Todiramphus tutus -- Wikipedia: Chattering kingfisher Todiramphus tutus atiu -- Wikipedia: Chattering kingfisher Todiramphus tutus -- Wikipedia: Chattering kingfisher Todiramphus tutus mauke -- Wikipedia: Chattering kingfisher Todiramphus veneratus -- Wikipedia: Society kingfisher Todiramphus veneratus youngi -- Wikipedia: Society kingfisher Todiramphus godeffroyi -- Wikipedia: Marquesan kingfisher Todiramphus chloris -- Wikipedia: Collared kingfisher Todiramphus chloris -- Wikipedia: Collared kingfisher Todiramphus chloris collaris -- Wikipedia: Collared kingfisher Todiramphus chloris laubmannianus -- Wikipedia: Collared kingfisher Todiramphus chloris teraokai -- Wikipedia: Collared kingfisher Todiramphus chloris humii -- Wikipedia: Collared kingfisher Todiramphus enigma -- Wikipedia: Talaud kingfisher Todiramphus albicilla -- Wikipedia: Mariana kingfisher Todiramphus albicilla -- Wikipedia: Mariana kingfisher Todiramphus albicilla -- Wikipedia: Mariana kingfisher Todiramphus albicilla -- Wikipedia: Mariana kingfisher Todiramphus albicilla -- Wikipedia: Mariana kingfisher Todiramphus chloris orii -- Wikipedia: Collared kingfisher Todiramphus saurophagus -- Wikipedia: Beach kingfisher Todiramphus saurophagus -- Wikipedia: Beach kingfisher Todiramphus chloris alberti -- Wikipedia: Collared kingfisher Todiramphus chloris nusae -- Wikipedia: Collared kingfisher Todiramphus chloris colcloughi -- Wikipedia: Collared kingfisher Todiramphus sordidus -- Wikipedia: Torresian kingfisher Todiramphus colonus -- Wikipedia: Islet kingfisher Todiramphus sanctus -- Wikipedia: Sacred kingfisher Todiramphus sanctus -- Wikipedia: Sacred kingfisher Todiramphus sanctus canacorum -- Wikipedia: Sacred kingfisher Todiramphus sanctus vagans -- Wikipedia: Sacred kingfisher Todiramphus chloris amoenus -- Wikipedia: Collared kingfisher Todiramphus sacer manuae -- Wikipedia: Pacific kingfisher Todiramphus sacer pealei -- Wikipedia: Pacific kingfisher Todiramphus chloris marinus -- Wikipedia: Collared kingfisher Todiramphus sanctus eximius -- Wikipedia: Sacred kingfisher Todiramphus sanctus vitiensis -- Wikipedia: Sacred kingfisher Todiramphus chloris ornatus -- Wikipedia: Collared kingfisher Todiramphus sacer -- Wikipedia: Pacific kingfisher Todiramphus chloris santoensis -- Wikipedia: Collared kingfisher Todiramphus chloris solomonis -- Wikipedia: Collared kingfisher Todiramphus pelewensis -- Wikipedia: Rusty-capped kingfisher Todiramphus pelewensis -- Wikipedia: Rusty-capped kingfisher Todiramphus pelewensis -- Wikipedia: Rusty-capped kingfisher Todiramphus reichenbachii -- Wikipedia: Pohnpei kingfisher Todiramphus cinnamominus -- Wikipedia: Guam kingfisher Todiramphus cinnamominus -- Wikipedia: Guam kingfisher Todiramphus cinnamominus -- Wikipedia: Guam kingfisher Todiramphus recurvirostris -- Wikipedia: Flat-billed kingfisher Todiramphus farquhari -- Wikipedia: Vanuatu kingfisher Todiramphus australasia -- Wikipedia: Cinnamon-banded kingfisher Todiramphus albonotatus -- Wikipedia: White-mantled kingfisher Todiramphus diops -- Wikipedia: Blue-and-white kingfisher Todiramphus macleayii -- Wikipedia: Forest kingfisher Todiramphus lazuli -- Wikipedia: Lazuli kingfisher Todiramphus leucopygius -- Wikipedia: Ultramarine kingfisher Todiramphus pyrrhopygius -- Wikipedia: Red-backed kingfisher Todiramphus funebris -- Wikipedia: Sombre kingfisher Todiramphus winchelli -- Wikipedia: Rufous-lored kingfisher Todiramphus nigrocyaneus -- Wikipedia: Blue-black kingfisher Syma megarhyncha -- Wikipedia: Mountain kingfisher Syma torotoro -- Wikipedia: Yellow-billed kingfisher Clytoceyx rex -- Wikipedia: Shovel-billed kookaburra Dacelo tyro -- Wikipedia: Spangled kookaburra Dacelo gaudichaud -- Wikipedia: Rufous-bellied kookaburra Dacelo leachii -- Wikipedia: Blue-winged kookaburra Dacelo leachii occidentalis -- Wikipedia: Blue-winged kookaburra Dacelo leachii cervina -- Wikipedia: Blue-winged kookaburra Dacelo leachii -- Wikipedia: Blue-winged kookaburra Dacelo leachii intermedia -- Wikipedia: Blue-winged kookaburra Dacelo novaeguineae -- Wikipedia: Laughing kookaburra Melidora macrorrhina -- Wikipedia: Hook-billed kingfisher Cittura cyanotis -- Wikipedia: Sulawesi lilac kingfisher Cittura cyanotis sanghirensis -- Wikipedia: Sulawesi lilac kingfisher Tanysiptera galatea -- Wikipedia: Common paradise kingfisher Tanysiptera hydrocharis -- Wikipedia: Little paradise-kingfisher Tanysiptera nigriceps -- Wikipedia: Black-capped paradise kingfisher Tanysiptera sylvia -- Wikipedia: Buff-breasted paradise kingfisher Halcyon albiventris -- Wikipedia: Brown-hooded kingfisher Halcyon albiventris orientalis -- Wikipedia: Brown-hooded kingfisher Halcyon cyanoventris -- Wikipedia: Javan kingfisher Halcyon gularis -- Wikipedia: Brown-breasted kingfisher Halcyon smyrnensis -- Wikipedia: White-throated kingfisher Halcyon pileata -- Wikipedia: Black-capped kingfisher Halcyon leucocephala -- Wikipedia: Grey-headed kingfisher Halcyon chelicuti -- Wikipedia: Striped kingfisher Halcyon chelicuti -- Wikipedia: Striped kingfisher Halcyon badia -- Wikipedia: Chocolate-backed kingfisher Halcyon coromanda -- Wikipedia: Ruddy kingfisher Halcyon coromanda linae -- Wikipedia: Ruddy kingfisher Halcyon coromanda major -- Wikipedia: Ruddy kingfisher Halcyon cyanoleuca -- Wikipedia: Woodland kingfisher Halcyon senegalensis -- Wikipedia: Woodland kingfisher Halcyon senegaloides -- Wikipedia: Mangrove kingfisher Halcyon malimbica -- Wikipedia: Blue-breasted kingfisher Pelargopsis capensis -- Wikipedia: Stork-billed kingfisher Lacedo pulchella -- Wikipedia: Banded kingfisher Lacedo pulchella melanops -- Wikipedia: Banded kingfisher Alcedo atthis -- Wikipedia: Common kingfisher Alcedo coerulescens -- Wikipedia: Cerulean kingfisher Alcedo hercules -- Wikipedia: Blyth's kingfisher Alcedo meninting -- Wikipedia: Blue-eared kingfisher Alcedo quadribrachys -- Wikipedia: Shining-blue kingfisher Alcedo semitorquata -- Wikipedia: Half-collared kingfisher Alcedo euryzona -- Wikipedia: Javan blue-banded kingfisher Ceyx argentatus -- Wikipedia: Southern silvery kingfisher Ceyx flumenicola -- Wikipedia: Northern silvery kingfisher Ceyx cyanopectus -- Wikipedia: Indigo-banded kingfisher Ceyx collectoris -- Wikipedia: New Georgia dwarf kingfisher Ceyx nigromaxilla -- Wikipedia: Guadalcanal dwarf kingfisher Ceyx sacerdotis -- Wikipedia: New Britain dwarf kingfisher Ceyx mulcatus -- Wikipedia: New Ireland dwarf kingfisher Ceyx solitarius -- Wikipedia: Papuan dwarf kingfisher Ceyx lepidus -- Wikipedia: Moluccan dwarf kingfisher Ceyx malaitae -- Wikipedia: Malaita dwarf kingfisher Ceyx melanurus -- Wikipedia: Philippine dwarf kingfisher Ceyx melanurus -- Wikipedia: Philippine dwarf kingfisher Ceyx melanurus samarensis -- Wikipedia: Philippine dwarf kingfisher Ceyx erithaca -- Wikipedia: Black-backed dwarf kingfisher Ceyx rufidorsa -- Wikipedia: Rufous-backed dwarf kingfisher Ceyx azureus -- Wikipedia: Azure kingfisher Ceyx websteri -- Wikipedia: Bismarck kingfisher Ceyx pusillus -- Wikipedia: Little kingfisher Ceyx fallax -- Wikipedia: Sulawesi dwarf kingfisher Corythornis cristatus -- Wikipedia: Malachite kingfisher Corythornis nais -- Wikipedia: Príncipe kingfisher Corythornis cristatus thomensis -- Wikipedia: Malachite kingfisher Corythornis vintsioides -- Wikipedia: Malagasy kingfisher Corythornis leucogaster -- Wikipedia: White-bellied kingfisher Corythornis madagascariensis -- Wikipedia: Madagascar pygmy kingfisher Ispidina lecontei -- Wikipedia: African dwarf kingfisher Ispidina picta -- Wikipedia: African pygmy kingfisher Ispidina picta natalensis -- Wikipedia: African pygmy kingfisher Ceryle rudis -- Wikipedia: Pied kingfisher Chloroceryle aenea -- Wikipedia: American pygmy kingfisher Chloroceryle inda -- Wikipedia: Green-and-rufous kingfisher Chloroceryle americana -- Wikipedia: Green kingfisher Chloroceryle amazona -- Wikipedia: Amazon kingfisher Megaceryle alcyon -- Wikipedia: Belted kingfisher Megaceryle torquata -- Wikipedia: Ringed kingfisher Megaceryle lugubris -- Wikipedia: Crested kingfisher Megaceryle maxima -- Wikipedia: Giant kingfisher Meropogon forsteni -- Wikipedia: Purple-bearded bee-eater Merops albicollis -- Wikipedia: White-throated bee-eater Todus angustirostris -- Wikipedia: Narrow-billed tody Momotus momota -- Wikipedia: Amazonian motmot SIHEK 3 individuals, 0.10–0.29% apart 2.02 points of nothing 0.29% to 2.30%, zero sequences
Radius = uncorrected ND2 p-distance from sihek MG008164, on a square-root scale, so an empty annulus covers page area in proportion to the distance interval it spans — the voids are honestly sized, not dramatised. Angle = tip order in the maximum-likelihood tree, so clades stay contiguous around the circle. Inked annuli are intervals containing no sequence at all. Hover a mark, or arrow-key through the chart, for its identity. The label stays for two seconds after you move away, so you can click the species name in it to open that bird on Wikipedia; the marks and the table names are links too. A dotted underline means the article covers the parent species, because the subspecies has none of its own.
0
sequences in the moat
2.02 pp
width of the void
2.30 %
nearest other kingfisher
0.29 %
spread within the sihek
7
empty annuli in the family
All 156 sequences and their ND2 distance from the sihek
SpeciesAccessionSubfamily Distance % Sites
Todiramphus cinnamominusMG008164Halcyoninae0.00001041
Todiramphus cinnamominusMG008165Halcyoninae0.19211041
Todiramphus cinnamominusKP291296Halcyoninae0.28821041
Todiramphus sacer pealeiKP291354Halcyoninae2.3041868
Todiramphus reichenbachiiMG008179Halcyoninae2.30551041
Todiramphus sacer manuaeKP291342Halcyoninae2.49761041
Todiramphus chloris santoensisKP291239Halcyoninae2.49761041
Todiramphus sacerKP291327Halcyoninae2.59371041
Todiramphus chloris amoenusKP291387Halcyoninae2.68971041
Todiramphus sanctus eximiusKP291298Halcyoninae2.68971041
Todiramphus sanctus vitiensisKP291386Halcyoninae2.68971041
Todiramphus chloris solomonisKP291365Halcyoninae2.68971041
Todiramphus colonusKP291383Halcyoninae2.78581041
Todiramphus recurvirostrisKP291389Halcyoninae2.83481023
Todiramphus chlorisKP291238Halcyoninae2.88181041
Todiramphus sanctus canacorumKP291326Halcyoninae2.88181041
Todiramphus sanctus vagansKP291369Halcyoninae2.88181041
Todiramphus chloris marinusKP291380Halcyoninae2.88181041
Todiramphus chloris ornatusKP291315Halcyoninae2.88181041
Todiramphus pelewensisKP291328Halcyoninae2.88181041
Todiramphus chlorisMG845643Halcyoninae2.97791041
Todiramphus chloris albertiKP291375Halcyoninae2.97791041
Todiramphus chloris nusaeKP291391Halcyoninae2.97791041
Todiramphus pelewensisKP291334Halcyoninae2.97791041
Todiramphus pelewensisKP291319Halcyoninae2.97791041
Todiramphus albicillaKP291351Halcyoninae3.07401041
Todiramphus albicillaKP291309Halcyoninae3.07401041
Todiramphus albicillaKP291295Halcyoninae3.07401041
Todiramphus albicillaKP291320Halcyoninae3.07401041
Todiramphus chloris colcloughiKP291390Halcyoninae3.07401041
Todiramphus sordidusKP291361Halcyoninae3.07401041
Todiramphus sanctusMG845661Halcyoninae3.07401041
Todiramphus albicillaKP291348Halcyoninae3.08881036
Todiramphus chloris oriiKP291370Halcyoninae3.17001041
Todiramphus sanctusKP291377Halcyoninae3.17001041
Todiramphus farquhariMG008171Halcyoninae3.17001041
Todiramphus saurophagusKP291374Halcyoninae3.26611041
Todiramphus saurophagusMG008181Halcyoninae3.26921040
Todiramphus ruficollarisMG008180Halcyoninae3.36221041
Todiramphus veneratusKP291261Halcyoninae3.36221041
Todiramphus veneratus youngiKP291371Halcyoninae3.36221041
Todiramphus godeffroyiKP291247Halcyoninae3.36221041
Todiramphus tutus atiuKP291302Halcyoninae3.45821041
Todiramphus tutusKP291382Halcyoninae3.45821041
Todiramphus tutus maukeKP291373Halcyoninae3.45821041
Todiramphus gertrudaeKP291321Halcyoninae3.55431041
Todiramphus tutusDQ111854Halcyoninae3.55431041
Todiramphus ruficollarisDQ111853Halcyoninae3.55771040
Todiramphus chloris laubmannianusKP291343Halcyoninae3.65031041
Todiramphus chloris collarisKP291384Halcyoninae3.74641041
Todiramphus chloris humiiKP291392Halcyoninae3.74641041
Todiramphus enigmaMG008168Halcyoninae3.74641041
Todiramphus chloris teraokaiKP291366Halcyoninae3.84251041
Todiramphus australasiaMG008163Halcyoninae5.66761041
Todiramphus diopsMG008167Halcyoninae6.0071849
Todiramphus albonotatusMG008161Halcyoninae6.53221041
Todiramphus macleayiiMG008176Halcyoninae6.62821041
Todiramphus lazuliMG008175Halcyoninae6.82041041
Todiramphus leucopygiusDQ111852Halcyoninae7.01251041
Todiramphus pyrrhopygiusMG008177Halcyoninae7.87701041
Todiramphus funebrisMG008173Halcyoninae8.0164973
Todiramphus nigrocyaneusKP291256Halcyoninae12.39191041
Todiramphus winchelliKP291381Halcyoninae12.48801041
Syma megarhynchaKP291344Halcyoninae13.83291041
Syma torotoroMG008149Halcyoninae14.12101041
Clytoceyx rexDQ111867Halcyoninae14.60131041
Actenoides lindsayiMG008067Halcyoninae14.69741041
Halcyon senegaloidesMG008131Halcyoninae14.98561041
Actenoides princepsMG008070Halcyoninae15.0943954
Actenoides concretusMG008065Halcyoninae15.36981041
Dacelo novaeguineaeMK116667Halcyoninae15.36981041
Actenoides monachusMG008068Halcyoninae15.4651860
Actenoides hombroniKP291331Halcyoninae15.46591041
Caridonax fulgidusMG008078Halcyoninae15.46591041
Dacelo leachii occidentalisMK116668Halcyoninae15.46591041
Dacelo leachii intermediaMK116666Halcyoninae15.46591041
Halcyon senegalensisMG008130Halcyoninae15.46591041
Dacelo gaudichaudMK116649Halcyoninae15.56201041
Dacelo leachiiDQ111865Halcyoninae15.56201041
Dacelo leachii cervinaMK116671Halcyoninae15.56201041
Dacelo leachiiMK116669Halcyoninae15.65801041
Halcyon cyanoleucaMK579324Halcyoninae15.74341029
Dacelo tyroMK116653Halcyoninae15.94621041
Halcyon malimbicaEF585389Halcyoninae16.13831041
Cittura cyanotisMG008101Halcyoninae16.16941039
Tanysiptera galateaMG008152Halcyoninae16.42651041
Halcyon leucocephalaMG008127Halcyoninae16.52261041
Halcyon badiaMG008117Halcyoninae16.52261041
Tanysiptera nigricepsMG008155Halcyoninae16.69961012
Melidora macrorrhinaMG008143Halcyoninae16.71471041
Cittura cyanotis sanghirensisMG008102Halcyoninae16.7421884
Actenoides bougainvillei excelsusMG008066Halcyoninae16.90681041
Halcyon gularisMG008124Halcyoninae16.92311040
Halcyon pileataMG008129Halcyoninae17.00291041
Tanysiptera sylviaMG008159Halcyoninae17.09891041
Halcyon smyrnensisMG008133Halcyoninae17.19501041
Halcyon albiventrisMG008116Halcyoninae17.48321041
Halcyon albiventris orientalisMK579323Halcyoninae17.57281030
Halcyon coromanda majorKF818925Halcyoninae17.60081017
Tanysiptera hydrocharisMG008154Halcyoninae17.67531041
Halcyon coromandaMG008121Halcyoninae17.67531041
Halcyon coromanda linaeKF818923Halcyoninae17.71651016
Halcyon cyanoventrisMG008122Halcyoninae17.7877669
Halcyon chelicutiMG008119Halcyoninae18.25171041
Halcyon chelicutiMK579322Halcyoninae18.29621033
Megaceryle alcyonEU166945Cerylinae18.73201041
Chloroceryle aeneaMG008096Cerylinae18.92411041
Chloroceryle indaMG008099Cerylinae19.11621041
Ceryle rudisMG008080Cerylinae19.21231041
Megaceryle lugubrisMG008140Cerylinae19.21231041
Chloroceryle amazonaMG008097Cerylinae19.30841041
Pelargopsis capensisMG008147Halcyoninae19.40441041
Alcedo herculesDQ640783Alcedininae19.50051041
Chloroceryle americanaMG008098Cerylinae19.59651041
Megaceryle maximaMG008141Cerylinae19.59651041
Megaceryle torquataMG008142Cerylinae19.69261041
Ceyx fallaxDQ640784Alcedininae20.07681041
Ceyx lepidusDQ111836Alcedininae20.17291041
Ceyx sacerdotisMG008093Alcedininae20.26901041
Ceyx malaitaeMG008085Alcedininae20.26901041
Ceyx melanurus samarensisMG008087Alcedininae20.28851040
Ceyx melanurusDQ640785Alcedininae20.36501041
Ceyx rufidorsaMN991393Alcedininae20.36501041
Ispidina leconteiEF585386Alcedininae20.36501041
Lacedo pulchellaMN991453Halcyoninae20.46111041
Ceyx mulcatusMG008089Alcedininae20.46111041
Ceyx solitariusMG008094Alcedininae20.46111041
Lacedo pulchella melanopsMG008137Halcyoninae20.55721041
Ceyx cyanopectusKC112606Alcedininae20.55721041
Ceyx azureusMG008081Alcedininae20.55721041
Ceyx websteriMG008095Alcedininae20.55721041
Corythornis madagascariensisMG008107Alcedininae20.55721041
Ceyx melanurusMG008086Alcedininae20.65321041
Corythornis vintsioidesMG008108Alcedininae20.65321041
Alcedo euryzonaMG008072Alcedininae20.74931041
Ispidina pictaEU166940Alcedininae20.76921040
Alcedo atthisEF585373Alcedininae20.84531041
Ceyx argentatusMH384618Alcedininae20.84531041
Ceyx flumenicolaMG008084Alcedininae20.84531041
Ceyx erithacaEF585385Alcedininae20.84531041
Ceyx pusillusMG008092Alcedininae20.84531041
Ceyx collectorisMG008082Alcedininae20.94141041
Ceyx nigromaxillaMG008090Alcedininae20.94141041
Corythornis leucogasterEF585379Alcedininae20.94141041
Ispidina picta natalensisMK579325Alcedininae21.04751031
Alcedo menintingMG008074Alcedininae21.13351041
Alcedo semitorquataDQ640781Alcedininae21.22961041
Corythornis naisEF585381Alcedininae21.32561041
Alcedo quadribrachysEF585382Alcedininae21.42171041
Corythornis cristatusEF585375Alcedininae21.51781041
Corythornis cristatus thomensisEF585384Alcedininae21.51781041
Alcedo coerulescensDQ640782Alcedininae22.37221037
Momotus momotaEU166942outgroup22.38231041
Todus angustirostrisDQ111829outgroup23.34291041
Merops albicollisEU021518outgroup25.93661041
Meropogon forsteniEU021535outgroup26.89721041

How to read the emptiness

The sihek's three sequenced individuals differ from each other by 0.10–0.29%. The nearest other kingfisher on Earth is 2.30% away. Between those two numbers lie 2.02 percentage points in which nothing exists — not one of the other 155 sequences, across every genus in the family.

That is the innermost ring. Ink the same way everywhere and the whole family turns out to be built of them: a second void at 3.84–5.67% walling off the core Todiramphus radiation, and a 4.38-point chasm at 8.02–12.39% separating the genus from every other kingfisher. Radiations leave gaps. This one happens to have a bird standing alone in the middle of it.

What the picture cannot tell you

A moat is not a measure of age, and this one is drawn from a single maternally inherited gene. ND2 traces the matriline only; introgression and incomplete lineage sorting are both plausible in an island archipelago, and a genome-scale study of the genus found that nuclear and mitochondrial data give no concordant topology at all.

Nor does the picture resolve which lineage the sihek split from. Its apparent sister on the tree, T. recurvirostris, carries only 66% bootstrap support. The isolation is solid; the ancestry is not.